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The soldier caste has anatomical and behavioural specialisations, and their only real purpose is to defend the colony. Many soldiers have large heads with highly modified strong jaws so enlarged they cannot feed themselves. Instead, such as juveniles, they are fed by employees.5556 Fontanelles, easy holes in the forehead which exude defensive secretions, are a characteristic of the family Rhinotermitidae.57 Many species are readily identified using the qualities of the soldiers' larger and darker head and large mandibles.53 Among certain termites, soldiers may utilize their globular (phragmotic) heads to obstruct their narrow tunnels.58 Different sorts of soldiers include minor and significant soldiers, and nasutes, which have a horn-like nozzle frontal projection (a nasus).53 These unique soldiers are able to spray noxious, sticky secretions containing diterpenes in their enemies.59 Nitrogen fixation has an important part in nasute nutrition.60.

The reproductive caste of a mature colony includes a fertile female and man, known as the queen and king.61 The queen of the colony is responsible for egg production for the colony. Unlike in ants, the king mates with her life.62 In some species, the abdomen of this queen swells up dramatically to increase fecundity, a characteristic known as physogastrism.61 Depending on the species, the queen starts producing reproductive winged alates at a certain period of year, and huge swarms emerge from the colony when nuptial flight begins.

A young termite nymph. Nymphs first moult into workers, but others might further moult to become soldiers or alates.

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Termites are often compared with the social Hymenoptera (ants and various species of bees and wasps), but their differing evolutionary origins result in significant differences in life span. In the eusocial Hymenoptera, the workers are entirely female. Males (drones) are haploid and develop from unfertilised eggs, while females (both workers and the queen) are both diploid and grow from fertilised eggs.

Depending on species, male and female workers may have different functions in a termite colony.63.

The entire life span of a termite begins with an egg, but is different from that of a bee or ant in that it goes through a developmental process called incomplete metamorphosis, with egg, nymph and adult stages.64 Nymphs resemble little adults, and go through a series of moults as they grow.

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The development of nymphs into adults can take months; the time frame depends on food availability, temperature, and the general population of this colony. Since nymphs are unable to feed themselves, employees must feed thembut workers also take part in the social life of their colony and also have certain different tasks to accomplish such as foraging, building or maintaining the nest or tending to the queen.5367 Pheromones regulate the caste system in termite colonies, preventing all but a very few of those termites from becoming fertile queens.68.

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Queens of the eusocial termite Reticulitermes speratus are capable of a long lifespan without sacrificing fecundity. These long-lived queens have a significantly lower degree of oxidative damage, including oxidative DNA damage, than workers, soldiers and nymphs.69 The lower degrees of harm appear to be due to increased catalase, an enzyme that protects against oxidative stress.69.

Termite alates only leave the colony when a nuptial flight takes place. Alate men and females pair up together and then land in search of a suitable spot for a colony.70 A termite king and queen do not mate until they find such a place. When they dothey excavate a chamber large enough for both, shut up the entrance and move to mate.70 After mating, the pair never go outdoors and spend the remainder of their lives in the nest.

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By way of instance, alates in certain species appear during the daytime in summer while others emerge during the winter.71 The nuptial flight may also begin at dusk, when the alates swarm around areas with lots of lights. The time when nuptial flight begins depends on the environmental conditions, the time of day, moisture, wind speed and precipitation.71 The number of termites in a colony also varies, with the larger species typically having 1001,000 individuals.

The queen only lays 1020 eggs in the very early phases of the colony, but places as many as 1,000 per day when the colony is several years old.53 At maturity, a main queen has a great capacity to lay eggs. In certain species, the adult queen has a greatly distended abdomen and may produce 40,000 eggs per day.72 Both mature ovaries may possess some here 2,000 ovarioles each.73 The abdomen increases the queen's body length to a number of times greater than before mating and reduces her ability to maneuver freely; attendant workers offer assistance.

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